Recall the formula for R-squared: \begin{equation} R^2=\frac{\sum\limits_{i=1}^N (\hat{Y_i}-\overline{Y})^2}{\sum\limits_{i=1}^N (\hat{Y_i}-\overline{Y})^2 + \sum\limits_{i=1}^N (\hat{Y_i}-Y_i)^2}= \frac{SS_{model}}{SS_{model}+SS_{residual}} \end{equation} In your first statistics class, you are taught that adding more regressors can not decrease R-squared, even if they have no relationship to the dependent variable.
This, however, does not apply to the constant term. At first pass, this seems hard to believe: An unconstrained model should always do at least as well as a constrained model.
The catch is, the variance explained by the constant term is not included in the calculation of R-squared - we subtract when calculating .
The no constant restriction implicitly sets to zero. This will increase both the model sum of squares and the residual sum of squares. The model sum of squares effect dominates, however, and is pushed towards one.
I discovered this today, regressing realized GDP on forecasted GDP. Although the sum of squared errors is nearly identical for both models, the model sum of squares is much larger for the no-intercept case:

fig

My takeaway: Be careful when writing your own regress command in Matlab, or any other language. Omitting a constant term can drastically change R-squared.